Tomatoes may get more attention, but throughout Italy, lemon also often finds its way into pasta.

Along the Amalfi coast, ring-shaped calamarata pasta is paired with clams, parsley, garlic and fried strips of a sweet local lemon. In Sicily, broken pasta comes studded with toasted pistachios, Parmesan, and a lemon’s juice and zest. And in Rome, the juice is whisked with olive oil, cream and Parmesan for al dente tagliatelle.

The sour and subtly sweet citrus balances the noodle’s starch and the sauce’s heft, functioning much like the dose of freshly ground black pepper that offsets pecorino Romano in cacio e pepe. And it serves much the same role as tomato often does, providing acidity that cuts through cheese.

In fact, the pairing of lemons and pasta is arguably more traditional than red sauce itself, as the fruit arrived in Italy at least 1,500 years before tomatoes.

Lemon features prominently in one pasta in our book “COOKish,” which limits recipes to just six ingredients without sacrificing flavor. Bright, bracing lemon zest and juice perk up the savoriness of garlic, pancetta and capers, while a dusting of golden crisp breadcrumbs adds texture.

This spaghetti also is a one-pot wonder: We cook the pasta in a minimal amount of water, and the starchy liquid that results forms a lightly thickened sauce that coats the noodles.

Lemon-Caper Spaghetti with Pancetta and Toasted Breadcrumbs Start to finish: 25 minutes Servings: 

4 to 6 1/2 cup panko breadcrumbs 

2 tablespoons extra-virgin olive oil 

Kosher salt and ground black pepper 

4 ounces pancetta, chopped 

4 medium garlic cloves, minced 

1 cup drained capers, chopped 

1 pound spaghetti

Grated zest and juice of 1 lemon In a large pot, cook the panko and oil, stirring, until golden brown. Transfer to a small bowl and stir in a pinch of salt. In the same pot, cook the pancetta, stirring, until crisp. Add the garlic and most of the capers; cook, stirring, until fragrant. Add 4 cups water and 1/2 teaspoon each salt and pepper, then boil. Stir in the pasta, cover and cook, stirring, until al dente. Off heat, stir in the remaining capers and lemon zest and juice; season with salt and pepper. Sprinkle with the breadcrumbs.

Optional garnish: Chopped fresh flat-leaf parsley or finely grated pecorino Romano cheese or both.